4/19/2023

Population Regulation

 In ecology, population regulation refers to the processes that control the growth and size of populations in a particular ecosystem. These processes can be influenced by both density-dependent and density-independent factors.

Density-dependent factors are those that become more important as a population becomes more crowded. For example, competition for limited resources such as food, water, and shelter, predation, and disease can all become more intense as a population grows. As a result, these factors can help to regulate the population size.

Density-independent factors, on the other hand, are those that affect a population regardless of its size. Examples of density-independent factors include natural disasters, such as floods or wildfires, changes in climate, and human activities such as habitat destruction and pollution.

In reality, population regulation is often a complex interplay between both density-dependent and density-independent factors. For example, a population may be growing rapidly due to favorable weather conditions (a density-independent factor) but may then experience a sudden increase in predation (a density-dependent factor) that slows or even reverses its growth.

Therefore, understanding the interplay between these factors is crucial for predicting and managing population growth and decline in real-life situations.

The Roles of Intraspecific Competition

 Intraspecific competition is the competition that occurs among individuals of the same species for resources like food, water, mates, and living space. The role of intraspecific competition can be significant in shaping the behavior, ecology, and evolution of a species.

Here are some of the roles of intraspecific competition:

  1. Regulating population size: Intraspecific competition can help regulate population size by limiting the number of individuals that can survive in a particular area. When resources are limited, individuals must compete for them, and some may not be able to access enough resources to survive.

  2. Driving natural selection: Intense intraspecific competition can lead to the evolution of traits that increase an individual's ability to compete for resources. Traits such as aggression, strength, and agility may become more common in a population as a result of this competition.

  3. Promoting dispersal: Intraspecific competition can also promote dispersal, where individuals move away from an area with high competition to find areas with more resources. This can lead to the formation of new populations and help maintain genetic diversity.

  4. Affecting social behavior: Intraspecific competition can influence social behavior within a species. For example, competition for mates can lead to the development of elaborate courtship displays, or to territorial behavior where individuals defend a particular area for exclusive use.

Overall, intraspecific competition plays an important role in shaping the ecology and evolution of a species. It can help regulate population size, drive natural selection, promote dispersal, and influence social behavior.

The two most basic models of population growth

 The two most basic models of population growth are the exponential growth model and the logistic growth model.

The exponential growth model assumes that populations grow without limits, with the rate of population growth being proportional to the current population size. Mathematically, this is described by the equation dN/dt = rN, where dN/dt is the rate of change in population size over time, N is the current population size, and r is the intrinsic rate of increase (i.e., the rate at which the population grows in the absence of any limiting factors). Exponential growth leads to a J-shaped curve, where the population increases rapidly at first and then accelerates further as the population size increases.

The logistic growth model, on the other hand, introduces limits to population growth, such as limited resources, space, or predation. The logistic growth equation is dN/dt = rN(1 - N/K), where K is the carrying capacity, or the maximum number of individuals that the environment can support. As the population approaches the carrying capacity, the growth rate slows down and eventually levels off, resulting in an S-shaped curve.

While neither model perfectly describes natural populations, they provide a useful starting point for understanding how populations can grow and what factors limit their growth. Population ecologists use these models to develop more complex models that incorporate other factors such as age structure, environmental variability, and density-dependent factors that can influence population growth and dynamics.

Survivorship curves

 Survivorship curves are a common tool used by population ecologists to understand how populations of organisms change over time. These curves graph the number of individuals surviving at different age intervals or life stages, typically from birth to death, and can be used to infer patterns of mortality and life expectancy within a population.

There are three main types of survivorship curves: Type I, Type II, and Type III, which differ in their shape and the age at which mortality rates increase.

  • Type I survivorship curves are characterized by high survivorship rates early in life, followed by a rapid decline in survivorship as individuals approach old age. This pattern is often seen in large mammals and other long-lived organisms with low infant mortality rates.


  • Type II survivorship curves show a roughly constant rate of survivorship across all age intervals. This pattern is seen in some birds, reptiles, and invertebrates.


  • Type III survivorship curves are characterized by low survivorship rates early in life, followed by a period of high survivorship in adulthood. This pattern is often seen in species that produce many offspring with little parental investment, such as insects and some fish.

By examining the survivorship curve of a population, ecologists can gain insights into factors that affect population growth and survival, such as predation, disease, and environmental conditions.

What is population size and density, and how can they be estimated?

 What is population size and density, and how can they be estimated?

Population size refers to the total number of individuals of a particular species living in a specific area. Population density is the number of individuals per unit area. The study of populations is called demography, and it provides mathematical tools to describe populations and investigate how they change.

The most accurate way to determine population size is to count all individuals within an area. However, this method is usually not feasible. Therefore, scientists sample a representative portion of each habitat and use this sample to estimate the population size and density.

For immobile organisms, such as plants or small, slow-moving organisms, a quadrat may be used. A quadrat is a square randomly located on the ground and used to count the number of individuals within its boundaries. For smaller mobile organisms, such as mammals, the mark and recapture method is often used. This method involves marking a sample of captured animals and releasing them back into the environment to mix with the rest of the population. Then, a new sample is captured, and scientists determine how many of the marked animals are in the new sample.

Species distribution pattern, which is the distribution of individuals within a habitat at a particular point in time, can provide further information about a population. Individuals within a population can be distributed randomly, clumped, or uniformly. Different distributions reflect important aspects of the biology of the species and affect the mathematical methods required to estimate population sizes.

Overall, demography is used to study the dynamics of a population, which involves understanding the changes in population size and density over time.

For mark and recapture method:

N = (M x n) / R Where:

  • N is the estimated population size
  • M is the number of individuals marked in the first capture
  • n is the total number of individuals captured in the second capture
  • R is the number of marked individuals recaptured in the second capture

For quadrat sampling method: Population size = (Total number of individuals in all quadrats / Area of one quadrat) x Total area of habitat

Density = Total number of individuals in all quadrats / Total area of habitat

Population Size and Density: A population is a group of organisms of the same species living in the same area. Population size is the total number of individuals in the population, while population density is the number of individuals per unit area. Population size and density can affect potential for adaptation and the interactions within a population such as competition for food and the ability of individuals to find a mate.

Estimating Population Size: The most accurate way to determine population size is to count all individuals within the area, but it is usually not feasible. Scientists usually sample a representative portion of each habitat to estimate the population as a whole. The methods used to sample populations vary depending on the characteristics of the organism being studied, such as using quadrats for immobile organisms or mark and recapture for smaller mobile organisms.

Species Distribution: A species distribution pattern is the distribution of individuals within a habitat at a particular point in time. Individuals within a population can be distributed at random, in groups, or equally spaced apart, known as random, clumped, and uniform distribution patterns, respectively. Different distributions reflect important aspects of the biology of the species.

Demography: Demography is the statistical study of population changes over time, including birth and death rates, immigration and emigration, and age structure. It is used to study the dynamics of a population.

Terrestrial Biomes

 Terrestrial Biomes

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Biomes are defined by characteristic temperatures and amounts of precipitation. The tropical rainforest is the most diverse of these biomes, found in equatorial regions with stable temperature and sunlight profiles, and high rainfall supporting rapid plant growth. The forest is characterized by vertical layering of vegetation, providing diverse habitats for the variety of plants, animals, and other organisms within the biome. The loss of leaves from taller trees during the dry season allows the growth of thick ground-level brush, which is absent in tropical rainforests. Extensive tropical dry forests occur in Africa, India, southern Mexico, and South America.


Terrestrial biomes refer to major ecological regions on land characterized by distinctive vegetation and animal communities. Aquatic biomes include both ocean and freshwater biomes, while terrestrial biomes are based on land.


Eight major terrestrial biomes

There are eight major terrestrial biomes: tropical rainforests, savannas, subtropical deserts, chaparral, temperate grasslands, temperate forests, boreal forests, and Arctic tundra.


Tropical rainforests

  • Equatorial regions.
  • The most diverse terrestrial biome.  
  • "evergreen," with year-round plant growth.
  • Stable temperature and sunlight profiles, with average temperatures ranging from 20°C to 34°C (68°F to 93°F)
  • High net primary productivity because the annual temperatures and precipitation values support rapid plant growth. 
  • High rainfall quickly leaches nutrients from the soils of these forests, which are typically low in nutrients.
  • Characterized by vertical layering of vegetation and the formation of distinct habitats for animals within each layer. 


Tropical dry forests

  • Characterized by a dry season of varying lengths and are found in Africa (including Madagascar), India, southern Mexico, and South America.
  • During the dry season, taller trees lose their leaves, opening up the canopy and allowing sunlight to reach the forest floor, allows for the growth of thick ground-level brush.


Savannas

  • Grasslands with scattered trees found in Africa, South America, and northern Australia
  • Temperatures averaging from 24°C –29°C (75°F –84°F)
  • Annual rainfall of 51–127 cm (20–50 in)
  • Extensive dry season and consequent fires
  • Dominated by grasses and forbs
  • Relatively few trees due to fire disturbance
  • Plants have well-developed root systems that allow them to re-sprout after a fire.


Subtropical Deserts

  • Exist between 15° and 30° north and south latitude and are centered on the Tropic of Cancer and the Tropic of Capricorn
  • Very dry with evaporation typically exceeding precipitation
  • Daytime soil surface temperatures above 60°C (140°F) and nighttime temperatures approaching 0°C (32°F)
  • Low annual precipitation of fewer than 30 cm (12 in) with little monthly variation and lack of predictability in rainfall
  • Low species diversity due to low and unpredictable precipitation
  • Desert species exhibit adaptations to conserve water
  • Most animal has adapted to a nocturnal life
  • North American deserts, Sahara Desert, Namib Desert in southwestern Africa


Cold Deserts

  • Deserts that experience freezing temperatures during the winter and any precipitation is in the form of snowfall
  • Gobi Desert in northern China, southern Mongolia, Taklimakan Desert in western China, Turkestan Desert, Great Basin Desert of the United States.


Chaparral

  • Also called scrub forest found in California, along the Mediterranean Sea, and along the southern coast of Australia
  • Annual rainfall ranging from 65 cm to 75 cm (25.6–29.5 in)
  • Summers are very dry with many plants dormant during this season
  • Dominated by shrubs and adapted to periodic fires
  • Some plants produce seeds that germinate only after a hot fire
  • Ashes left behind after a fire are rich in nutrients like nitrogen that fertilize the soil and promote plant regrowth
  • Fires are a natural part of the maintenance of this biome and frequently threaten human habitation.


Temperate Grasslands

  • Central North America, known as prairies, and in Eurasia, known as steppes
  • Pronounced annual fluctuations in temperature with hot summers and cold winters
  • Dense vegetation and fertile soils due to the subsurface of the soil being packed with roots and rhizomes

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